I thought I was onto something special with this distinctively marked fly. There was nothing too hard about the route through the family key which brought me to a new family, and even better, it had very few genera and species which meant that I might be able to identify it completely. A poster fly for ease of identification, an easy acalypterate, a fly so full of character that it has a cryptic message written in fuzzy ink across its wings – nu on the left and un on the right – nu-un.
But when I went through online keys and checked out photos I was clearly on the wrong track – the best fit was a completely different looking beast with WaaW written across its wings, which is pretty much how I felt. Instead of being from an exciting new family, I must have gone wrong somewhere in the key and it was probably one of those spotty-winged tephritids. Except that none of the tephritid wing patterns matched, and the key character for the family, the Sc vein being bent at a right angle and fainter as it reaches the wing margin was not right (Sc is the thickish vein that curves up and just makes it into the top of the white part of the black U shape).
It turns out that the fly is a dead ringer for Herina frondescentiae, a member of the family Ulidiidae. Going through the key again, there should be two breaks in the vein that runs along the top of the wing. You can see one at the extreme left side of the wing, and there should be another roughly where the Sc vein meets the wing edge – but there is no sign any break there. Everything else about the fly is right – including its reddish face with silvery side patches and “distinctive” nu-un message – and although it wasn’t the family I first thought it might be (Platystomatidae), it’s still a new family – a new ‘un in fact!
New families are now getting harder to find – I had high hopes from a batch caught during a work day lunchtime break with a fellow dipterist. He pointed out a mob of flies purposefully cruising back and forward over a stream, just out of net reach until the wind eddied slightly and brought them within striking distance. They sported unusually inflated front feet, but turned out to be Hilara, a genus of Empididae. Also promising was a fly with jauntily spotted wings swept from the edge of a bean field, but this was Trypeta zoe, a differently patterned Tephritid. Next, a greenish metallic fly swept from a buttercup-filled meadow was at first sight a hover fly, but seemed to lack the distinctive spurious vein of the Syrphidae. But all the other routes through the key ended up at families that were plainly wrong, and so eventually I concluded that it was Lejogaster metallina, a hover fly that can be confused with a soldier fly (Stratiomyidae); unfortunately for me this was the hover fly. My last hopes were pinned on a tiny fly also swept from the meadow that I thought might have been Drosophilidae, but my companion knew better, being an expert on the family. Of course he was right, and it turned out to be another Piophilidae.
Finding new families was becoming hard work, so it was somewhat of a relief when, as I was enjoying my breakfast outside, my toes dipped in clover, a distinctively purposeful and noisy fly joined me in the sunshine. As I got up to fetch my net it flitted away, but since we had both identified the sunniest and most sheltered spot in the garden, I suspected it might be back. With the net by my side I continued to contemplate the possibilities of the day. Half a cup of coffee later it landed on my net and with an expert flick of the wrist I curtailed its morning salutations. This was, without my needing to go through the key, a flesh fly of the family Sarcophagidae, as smartly turned out as a chap heading out to work in suit and tie. Its thorax was a neat pinstripe of grey and black stripes, set off nicely by gold dusting between the eyes, the abdomen tesselated in glinting black and grey, an eye-catching tweed.
I assumed that being such a large beast, taking the identification further would be trivial, possibly not even needing the microscope. I started off using my photocopied key (1954) to Calliphoridae by van Emden which includes what was then the subfamily Sarcophaginae. Finding the genus was a struggle through couplets asking if the hind coxae was setulose, about a row of inclinate frontal setae, and if the mid tibia had a postmedian ventral seta, but I cautiously arrived at Sarcophaga, for which the key noted: “The chaetotaxctic characters [ie. the bristles] are somewhat unreliable and the only way of obtaining quite definite identifications is by comparing the male genitalia.” With an updated version of the genus key I stumbled along, confirming that vein R1 was not setulose, that there were four presutural dorsocentrals, that abdominal tergite 3 had strong marginals – which brought me, possibly, to a choice of three widespread species, S. subvicina which is a parasite of earthworms, or one of a seemingly indistinguishable pair – S. carnaria and S. variegata – for which the first of which the van Emden key notes: “Larvae in decomposing organic matter, including dead animals, sometimes parasitic, even in man, causing intestinal myiasis etc.” The “etc.” was slightly worrying, as was the thought that worm or man, quick or dead, there was flesh fly for whom you were the right kind of food.
Psychologists know that a good way of getting people to talk about things that matters to them is to get them to talk about something else. What did you dream about last night? How did you feel about that? What associations does that image bring up? Or at another level of distraction, there is the ink-spot test where a meaningless blog (Freudian slip there – I meant blob) becomes a particular something that can be talked about as if it were meaningless.
Well have a look at this fly wing and tell me what you see. Relax – there is no right answer. Personally, I see Santa waving a greeting from his toy-laden sledge as a copiously maned lion pulls him across an icy lake while frenzied onlooking snow ghouls attack from every direction. Fascinating. Santa being Father Christmas of course, and the pull of lying (lion) might be man-ed? Go on. Well, this is the picturesque wing of Tephrita neesia, a member of the family Tephritidae, and a member of the superfamily Tephritoidea, commonly known, although I suspect by very few people. as picture-winged flies. Superfamily – interesting. How do you picture yourself in your family?
As the fly expert, obviously. It so happens that I have an unopened identification key to British Tephritidae on my shelves, Royal Entomological Society, 1988 by I.M. White. Just say that name again would you – I. M. White – I am right. Tephritidae – too afraid today. Would you say that you worry about, let’s say mistaking one fly family for another? Getting the identification wrong, so the book, the identity was unopened, unexplored…. Instead of the usual worries about the position of bristles on the face or how one wing vein meets another, the Tephritidae key is almost entirely based on the pattern of dark splodges, or by inverting the image, where the wing is clear or hyaline as it more obscurely terms it. You mentioned “inversion” and looking for bristles on a face – does either of these evoke a memory from your childhood perhaps?
The host for this fly is the ox eye daisy, which happens to be one of the flowers in the seed mix which I strewed upon what used to be the boys’ football pitch and is now the Meadow and where in late summer I can be glimpsed, topless wielding a scythe like the grim reaper himself. Wonderful! This is very interesting material indeed. Same time again next week – and just bring along whatever you catch – I think this blog of yours is going to be very fruitful material for us.